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Behavior

Free-ranging cats are active both day and night, although they tend to be slightly more active at night. The timing of cats' activity is quite flexible and varied, which means that house cats may be more active in the morning and evening (crepuscular behavior), as a response to greater human activity at these times. House cats have territories that vary quite a lot in size, in one study ranging from seven to 28 hectares. Although they spend the majority of their time in the vicinity of their home, they can range many hundreds of meters from this central point.

Cats conserve energy by sleeping more than most animals, especially as they grow older. The daily duration of sleep varies, usually 12–16 hours, with 13–14 being the average. Some cats can sleep as much as 20 hours in a 24-hour period. The term cat nap refers to the cat's ability to fall asleep (lightly) for a brief period and has entered the English lexicon—someone who nods off for a few minutes is said to be "taking a cat nap".

Sociability

Cats are a social species, and free-living cats tend to form feral cat colonies if there is sufficient food, which are based around groups of co-operating females. Within this group one cat is usually dominant over the others. Each cat in a colony holds a distinct territory, with sexually active males having the largest territories, which are about ten times larger than those of female cats and may overlap with several females' territories. These territories are marked by urine spraying, by rubbing objects at head height with secretions from facial glands and by defecation. Between these territories are neutral areas where cats watch and greet one another without territorial conflicts. Outside these neutral areas, territory holders usually chase away stranger cats, at first by staring, hissing, and growling, and if that does not work, by short but noisy and violent attacks. Despite some cats cohabiting in colonies, cats do not have a social survival strategy, or a pack mentality and always hunt alone.

As part of cat sociability they use a variety of vocalizations for communication, including meowing, purring, "trilling", hissing, growling, squeaking, chirping, clicking, and grunting.Their types of body language: position of ears and tail, relaxation of whole body, kneading of paws, are all indicators of mood. The tail and ears are particularly important social signal in cats, with a raised tail acting as a friendly greeting. Tail raising also indicates the cat's position in the group's social hierarchy, with dominant individuals raising their tails less often than subordinate animals. Nose-touching is also a common greeting and may be followed by social grooming, which is solicited by one of the cats raising and tilting its head. However, some pet cats are poorly socialized. In particular older cats may show aggressiveness towards newly-arrived kittens, which may include biting and scratching; this type of behavior is known as Feline Asocial Aggression.

For cats, life in proximity with humans (and other animals kept by humans) amounts to a "symbiotic social adaptation". They may express great affection towards their human companions, especially if they imprint on them at a very young age and are treated with consistent affection. It has been suggested that, ethologically, the human keeper of a cat functions as a sort of surrogate for the cat's mother, and that adult domestic cats live their lives in a kind of extended kittenhood, a form of behavioral neoteny.

Grooming

Cats are known for their fastidious cleanliness, spending many hours licking their coats. The cat's tongue has backwards-facing spines about 500 micrometres long, which are called papillae. These are quite rigid as they contain keratin. These spines allow cats to groom themselves by licking their fur, with the rows of papillae acting like a hairbrush. Some cats, particularly longhaired cats, occasionally regurgitate hairballs of fur that has collected in their stomachs from grooming. These clumps of fur are usually sausage-shaped and about two to three centimeters long. Hairballs can be prevented with remedies that ease elimination of the hair through the gut, as well as regular grooming of the coat with a comb or stiff brush.


The hooked papillae on a cat tongue act like a hairbrush to help clean and detangle fur.

Fighting

In domestic cats, males are more likely to fight than females. In feral cats, the most common reason for cat fighting is when two males are competing to mate with a female: here most fights will be won by the heavier male. Another possible reason for fighting in domestic cats is when the cats have difficulties in establishing a territory within a small home. Female cats will also fight over territory or to defend their kittens. Spaying females and neutering males will decrease or eliminate this behavior in many cases.


Cats intimidate opponents by arching their backs, raising their fur, turning sideways, and hissing.

Fighting cats make themselves appear more impressive and threatening by raising their fur and arching their backs, thus increasing their apparent size. Attacks usually comprise powerful slaps to the face and body with the forepaws as well as bites, but serious damage is rare; usually the loser runs away with little more than a few scratches to the face, and perhaps the ears. Cats will also throw themselves to the ground in a defensive posture to rake their opponent's belly with their powerful hind legs.

Normally, serious injuries from fighting will be limited to infections of scratches and bites, though these can occasionally kill cats if untreated. In addition, bites are probably the main route of transmission of feline immunodeficiency virus (FIV). Sexually active males will usually be in many fights during their lives, and often have decidedly battered faces with obvious scars and cuts to the ears and nose.

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